Key to Intertidal Aleocharine Genera of
America North of Mexico
(Coleoptera: Staphylinidae)
Kee-Jeong Ahn
(This "Key to Intertidal Aleocharine Genera
of America North of Mexico" is an
UNREVIEWED DRAFT provided for INFORMATION ONLY. It is NOT PUBLISHED,
and should not be cited, quoted or referenced in published papers.)
July 1996
*The mouthparts need to be dissected in order to see the mouthpart characters
referred to in these keys.
Adults
1. Front tarsus with 5 articles; maxillary palpi and labial palpi with
pseudo-articles (Figure)
---------------------------------------------------------------------------
Aleochara
Front tarsus with 4 articles; mxillary palpi and labial palpi without
pseudo-articles ------ 2
2. Maxillary palpi extremely elongate; labial palpi stylate -------------------------
3
Maxillary palpi not elongate; labial palpi not stylate -------------------------------------
6
3. Galea with long seate on mesal surface; lacinia without distinctive
distribution of spines
interdigitating with spinose scales; anterior-lateral angles of mentum
not prolonged into spinose
processes; gland opeing on tergite VII large ----------------------------------
Diglotta
Galea without setae on mesal surface (Figure);
lacinia with distinctive distribution of spines
interdigitating with spinose scales (Figure);
anterior-lateral angles of mentum prolonged into spinose
processes (Figure);
gland opening on tergite VII small ---------------------------------------
4
4. Body fusiform; hypomera not visible in lateral view; postero-lateral
margins of elytra
sinuate; tergite X bifid -------------------------------------------------------
Myllaena
Body not fusiform; hypomera visible in lateral view; postero-lateral
margins of elytra not
sinuate; tergite X not bifid --------------------------------------------------------------------5
5. Pronotum subquadrate; mesocoxal cavities contiguous; posterior margin
of mesocoxal
acetabula not margined; mesosternal process sharply pointed; abdominal
segment IX without
distinct circular setal pattern; tergites impressed at base --------------------
Bryothinusa
Pronotum transverse; mesocoxal cavities widely separated; posterior
margin of mesocoxal
acetabula margined; mesosternal process broadly rounded; abdominal
segment IX with distinct
circular setal pattern (Figure);
tergites not impressed at base --------------------------- Rothium
6. Mandible with blunt apex (Figure);
front and middle tibiae with spines (Figure)
------------- 7
Mandible with sharp apex; front and middle tibiae without spines ------------------------
9
7. Head without infraorbital carina; elytra longer than pronotum; hind
wings present; internal
circular sclerites present at base of metasternum ------------------------------Tarphiota
Head with infraorbital carina; elytra shorter than pronotum; hind wings
absent; internal
circular-shaped sclerites absent at base of metasternum ------------------------------------
8
8. Antennomeres 5--10 transverse; mandible (Figure)
with subapically
oriented median tooth; ligula (Figure)
narrow and bifid; scutellum octagonal; mesocoxal
cavities contiguous; mesosternal process sharp; elytra slightly sinuate
apico-laterally; tarsal
formula 4-4-5 -----------------------------------------------------------------
Thinusa
Antennomeres 8--10 transverse; mandible (Figure)
with laterally oriented median tooth;
ligula wide and bifid; scutellum diamond-shape; mesocoxal cavities
separated;
mesosternal process round; elytra strongly sinuate apico-laterally;
tarsal formula 4-5-5
---------------------------------------------------------------------------
Pontomalota
9. Middle tarsus with 5 articles ----------------------------------------------
Atheta
Middle tarsus with 4 articles ---------------------------------------------------------
10
10. Base of head and pronotum longitudinally impressed; neck present;
pronotum narrowed
behind middle ----------------------------------------------------------------
Bryobiota
Base of head and pronotum not impressed; neck absent; pronotum not narrowed
behind
middle -------------------------------------------------------------------------------
11
11. Mesocoxa separated ----------------------------------------------------------12
Mesocoxa contiguous ------------------------------------------------------------
13
12. Ligula bifid; infraorbital carina absent; tergite III-V impressed
at base
------------------------------------------------------------------------------
Thinobiosus
Ligula simple; infraorbital carina present; tergite III-VI impressed
at base
---------------------------------------------------------------------------------
Heterota
13. Ligula bifid; elytra longer than pronotum; hind wings present --------------------------
14
Ligula simple; elytra shorter than pronotum; hind wings absent ----------------------------
15
14. Spinule-like vestige absent on galeal surface; two medial setae
present on prementum;
setae absent on ligula; hypoglossa not distinct ------------------------------------
Cameronium
Spinule-like vestige present on galeal surface; one medial seta present
on prementum; setae
present on ligula; hypoglossa distinct, with long spines (Figure)
--------------------- Salinamexus
15. Median tooth of right mandible (Figure)
not triangular; lacinia (Figure)
distinctive, with large apical spine, four short spine more medially
and one large spine behind
these; anterior margin of abdominal tergites III--VI deeply and broadly
V-shaped; apical lobe
of paramere (Figure)
more or less diamond-shaped ------------------------------ Amblopusa
Median tooth of right mandible triangular; lacinia with about 10 spines
of subequal size in a row
near apex anterior margin of abdominal tergites III--VI straight; apical
lobe of paramere (Figure)
long ------------------------------------------------------------------------------------
16
16. Labrum (Figure)
transverse; mandible (Figure)
with small teeth between apex and median
tooth; article 3 of maxillary palpus ovoid; several setae only on mesal
surface of galea,
and apex of galea with setae (Figure);
mentum (Figure) triangular,
deeply incised with
V-shaped emargination at apex; ratio of pronotal length to elytral
length less than 1.3;
metasternal length almost same as mesocoxal width; posterior margin
of abdominal tergite X not
sclerotized; spermathecal tube (Figure)
long and coiled ------------------- Paramblopusa
Labrum semicircular or narrow trapezpidal; article 3 of maxillary palpus
dilated distally;
many setae on mesal surface of galea, and apex of galea with dense
setae; mentum not triangular;
mandible without or with large teeth between apex and median tooth;
ratio of
pronotal length to elytral length more than 1.3; metasternal length
shorter than half of
mesocoxal width; posterior margin of abdominal tergite X sclerotized;
spermathecal tube
short and straight --------------------------------------------------------------------------
17
17. Setae absent between facets of compound eyes; labrum (Figure)
very narrow, trapezoidal
and more than 50 setae present; mandible (Figure)
with about 5 large teeth between
apex and median tooth, and not curved ventrally; lacinial setae and
galeal setae (Figure)
distributed
uniformly over surface; mentum not small and anterior margin not deeply
emarginate; tarsus
with spatulate setae; female abdominal sternite VIII prolonged posteriorly;
abdominal
tergites not impressed -------------------------------------------------Liparocephalus
Setae present between facets of compound eyes; labrum (Figure)
more or less semicircular
and fewer than 30 setae present; mandible (Figure)
without large teeth between apex and median
tooth, and curved ventrally; lacinial setae and galeal setae (Figure)
distributed on mesal surface;
mentum small, its anterior margin deeply emarginate; tarsus without
spatulate setae; female
abdominal sternite VIII not prolonged posteriorly; abdominal tergites
III-VI impressed
---------------------------------------------------------------------------Diaulota
Late Instar Larvae
1. Anterior margin of labrum (Figure)
with distinctive denticles; mala (Figure)
with about 20 spinose
setae on mesal surface; ligula broad (Figure)
-------------------------------------------------------- 2
Anterior margin of labrum without distinctive denticles; mala with less
than 10 spinose setae on
mesal surface; ligula narrow -----------------------------------------------------------------------
3
2. Article 1 of antennae (Figure)
longer than wide; sensory appendage about as long as article 3;
mandible (Figure)
with median tooth large, followed by 3 large serrations and more or less
round serration
distinctively present -----------------------------------------------------------------------
Rothium
Article 1 of antennae wider than long; sensory appendage about 2 times
as long as article 3;
mandible with median tooth large, followed by 4 large serrations and
more or less round serration absent
-------------------------------------------------------------------------------------------
Bryothinusa
3. Sensilla IIS3 and IIIS4 of antennae absent; mandible without serration
between apex and median
tooth; 3--5 pointed spines on anterolateral margin of labium (Figure);
submentum narrow and unsclerotized;
1 robust spine on tarsus (Figure);
indistinctly articulated urogomphus; 4 large sclerotized anal hooks
(Figure) present
---------------------------------------------------------------------------
Amblopusa
Sensilla IIS3 and IIIS4 of antennae (Figure)
present; mandible (Figure)
with serration between apex
and median tooth; 1 pointed spine or 2--3 more or less sword-shaped
spines on anterolateral margin of
labium; submentum broad and sclerotized; 2 robust spines on tarsus;
urogomphus articulated or fused to main
body; 4 small unsclerotized anal hooks present or absent --------------------------------------
4
4. Ratio of length of sensory appendage to length of article 3 of antenna
(Figure) less than
1.0;
mandible (Figure)
with large serrations between apex and median tooth; 1 pointed spine present
on
antero-lateral margin of labium (Figure);
urogomphus (Figure)
articulated; long, slender seta present
on apex of urogomphus; 4 small unsclerotized anal hooks present ---------------------
Liparocephalus
Ratio of length of sensory appendage to length of article 3 of antenna (Figure)
more than 1.0;
mandible (Figure)
with small serrations between apex and median tooth; 2--3 more or less
sword-shaped
spines on anterolateral margin of labium (Figure);
urogomphus (Figure)
fused to main body; without long,
slender apical seta on urogomphus; anal hooks absent --------------------------------------
Diaulota
Scales of line drawings = 0.1mm.
|