A catalogue of the athetine species of America north of Mexico (Coleoptera: Staphylinidae: Aleocharinae: Athetini)

By Vladimir I. Gusarov

Last updated: December 15, 2003

This list is the result of my revision of the types of North American athetines. Most of the species were described by Casey (Casey 1885, 1893, 1906, 1910, 1911) about 100 years ago and have never been revised since that time. To separate species, Casey relied exclusively on external characters and often described the same species several (up to 14!) times (Gusarov 2003f). My study allowed to discover numerous synonyms among the species described by Casey and other authors. These synonyms are listed below, but for many genera the results have not been published yet.

Most Nearctic species of athetines belong to the same genera as the species distributed in the Palaearctic region. However before my study many Nearctic species were not correctly assigned to genera. Casey created numerous generic and subgeneric names, and the relation of these taxa to Palaearctic genera has not been clarified. One of the results of my type revision is the correct assignment of Nearctic species to genera. Since this is an on-going project, the list is still preliminary. For the time being I prefer to assign the species to large genera (for example, Atheta sensu lato) without subgeneric division.

Some of the species described in Athetini should be moved to other tribes. Please see the list of removed species below the Athetini catalogue.

The references are listed at the end of the list.

Generic names and species names within genera are arranged alphabetically. To search for a name you can use your browser’s search function (Ctrl+F on PC; Apple+F on Mac).

If you use this catalogue in your work please cite it in your references. This web page can be cited as follows:

Gusarov, V.I. (2001-2003) A catalogue of the athetine species of America north of Mexico (Coleoptera: Staphylinidae: Aleocharinae: Athetini). <http://nhm.ku.edu/ksem/peet/cataweb.htm>. Last updated: December 15, 2003. Date of last access: MM/DD/YYYY.

This project is supported by the National Science Foundation PEET grant DEB-9978110 to Steve Ashe.

Tribe Athetini Casey, 1910

Genera	Valid species names	Synonyms	The genera (with subgenera) in which the species were originally described	Page num-bers in the ori-ginal publi-cations	The type locality states or provinces	Recent publications with diagnoses (for valid species) or confirmation of synonymy (for synonyms)
Acrotona	absona Casey, 1910		Acrotona (Engamota Casey, 1910) Thomson, 1859	152	CA
Acrotona	acutella Casey, 1910		Colpodota Mulsant & Rey, 1873	156	NY
		acutella simulata Casey, 1910	Colpodota Mulsant & Rey, 1873	156	MO
		sobria Casey, 1910	Colpodota Mulsant & Rey, 1873	157	MS
Acrotona	assecla Casey, 1910		Colpodota Mulsant & Rey, 1873	158	NY
Acrotona	austiniana Casey, 1910		Colpodota Mulsant & Rey, 1873	158	TX
Acrotona	cupiens Casey, 1910		Dolosota (s.str.) Casey, 1910	140	NY
		comis Casey, 1910	Dolosota (s.str.) Casey, 1910	140	IA
Acrotona	curata Casey, 1910		Sableta (Anatheta Casey, 1910) Casey, 1910	112	VA
Acrotona	dilutipennis Motschulsky, 1858		Homalota Mannerheim, 1830	252	Oriental Region
Acrotona	discreta Casey, 1893		Eurypronota Casey, 1893	335	IA
Acrotona	egens Casey, 1910		Colpodota Mulsant & Rey, 1873	160	CA
Acrotona	egregiella Casey, 1910		Colpodota Mulsant & Rey, 1873	156	NY
		acuminata Casey, 1910	Colpodota Mulsant & Rey, 1873	155	RI
		defessa Casey, 1910	Colpodota Mulsant & Rey, 1873	158	RI
		insulsa Casey, 1910	Colpodota Mulsant & Rey, 1873	155	RI
		sequestralis Casey, 1910	Colpodota Mulsant & Rey, 1873	159	IA
Acrotona	fatigans Casey, 1910		Colpodota Mulsant & Rey, 1873	161	CA
Acrotona	fontinalis Casey, 1910		Dolosota (Reania Casey, 1910) Casey, 1910	146	CO
Acrotona	hebeticornis Notman, 1920		Acrotona (s. str.) Thomson, 1859	730	FL
Acrotona	luteola Erichson, 1839		Homalota Mannerheim, 1830	114	North America
		alumna Casey, 1910	Dolosota (s.str.) Casey, 1910	142	MO
		collaris Casey, 1906	Pancota Casey, 1906	346	NY
		euphonia Casey, 1910	Dolosota (s.str.) Casey, 1910	143	NY
		lacertina Casey, 1910	Dolosota (s.str.) Casey, 1910	141	RI
		panda Casey, 1910	Dolosota (s.str.) Casey, 1910	143	DC
		redundans Casey, 1910	Dolosota (s.str.) Casey, 1910	137	NY
		restricta Casey, 1910	Dolosota (s.str.) Casey, 1910	142	MS
		serva Casey, 1910	Dolosota (s.str.) Casey, 1910	144	MA
		sophista Casey, 1910	Dolosota (s.str.) Casey, 1910	143	IA
Acrotona	obliquata Casey, 1910		Strigota Casey, 1910	183	CA
Acrotona	onthophila Lohse in Lohse et al., 1990		Acrotona Thomson, 1859	198	YT	Lohse et al. 1990
Acrotona	opica Casey, 1911		Oxypoda Mannerheim, 1830	46	CA
Acrotona	pasadenae Bernhauer, 1906		Atheta (Acrotona Thomson, 1858) Thomson, 1859	338	CA
		vaniuscula Casey, 1911	Dimetrota (Dimetrotina Casey, 1911) Mulsant & Rey, 1873	144	NV
Acrotona	petulans Casey, 1910		Dolosota (Microlia Casey, 1910) Casey, 1910	145	NJ
Acrotona	picescens Notman, 1920		Acrotona (s. str.) Thomson, 1859	729	FL
Acrotona	reclusa Casey, 1910		Dolosota (Aremia Casey, 1910) Casey, 1910	146	NY
Acrotona	recondita Erichson, 1839		Homalota Mannerheim, 1830	123	PA
		apacheella Casey, 1910	Arisota Casey, 1910	135	AZ
		insueta Casey, 1910	Arisota Casey, 1910	134	RI
		pomonensis Casey, 1910	Arisota Casey, 1910	135	CA
		speculifer Casey, 1910	Arisota Casey, 1910	135	CA
		tetricula Casey, 1910	Arisota Casey, 1910	134	NY
		umbrina Casey, 1910	Arisota Casey, 1910	136	CA
Acrotona	scopula Casey, 1893		Eurypronota Casey, 1893	335	RI
		abundans Casey, 1910	Dolosota (s.str.) Casey, 1910	139	MO
		flaccida Casey, 1910	Dolosota (s.str.) Casey, 1910	139	MS
		laetabilis Casey, 1911	Pancota Casey, 1906	154	NY
		redundans tergina Casey, 1910	Dolosota (s.str.) Casey, 1910	138	PA
		secunda Casey, 1910	Dolosota (s.str.) Casey, 1910	138	IA
		sequax Casey, 1910	Dolosota (s.str.) Casey, 1910	138	IA
Acrotona	smithi Casey, 1910		Coprothassa Thomson, 1859	166	NY
Acrotona	sonomana Casey, 1910		Colpodota Mulsant & Rey, 1873	162	CA	Gusarov 2003g
		abdicans Casey, 1910	Colpodota Mulsant & Rey, 1873	163	CA	Gusarov 2003g
		inceptor Casey, 1910	Colpodota Mulsant & Rey, 1873	162	CA	Gusarov 2003g
		laxella Casey, 1910	Colpodota Mulsant & Rey, 1873	164	CA	Gusarov 2003g
		pupilla Casey, 1911	Colpodota Mulsant & Rey, 1873	155	CA	Gusarov 2003g
		repentina Casey, 1910	Colpodota Mulsant & Rey, 1873	163	CA	Gusarov 2003g
		seclusa Casey, 1911	Strigota (Eustrigota Casey, 1911) Casey, 1910	165	CA	Gusarov 2003g
Acrotona	subpygmaea Bernhauer, 1909		Atheta (Acrotona Thomson, 1858) Thomson, 1859	526	MA
		avia Casey, 1910	Colpodota Mulsant & Rey, 1873	154	RI
		puritana Casey, 1910	Colpodota Mulsant & Rey, 1873	154	MA
Acrotona	trossula Casey, 1910		Colpodota Mulsant & Rey, 1873	159	AZ
Acrotona	turbans Casey, 1910		Colpodota Mulsant & Rey, 1873	157	TX
Acrotona	vafra Casey, 1910		Dolosota (s.str.) Casey, 1910	140	NC
Acrotona	zephyrina Casey, 1910		Colpodota Mulsant & Rey, 1873	160	CA
		torvula Casey, 1910	Colpodota Mulsant & Rey, 1873	162	CA
Adota	gnypetoides Casey, 1910		Atheta (Adota Casey, 1910) Thomson, 1858	69	CA	Gusarov 2003e
Adota	maritima Mannerheim, 1843		Homalota Mannerheim, 1830	224	AK	Gusarov 2003e
		immigrans Easton, 1971	Atheta (Phyconoma Easton, 1971) Thomson, 1858	25	Palaearctic Region	Gusarov 2003g
		insons Casey, 1911	Atheta (Adota Casey, 1910) Thomson, 1858	125	CA	Gusarov 2003e
		massetensis Casey, 1910	Atheta (Adota Casey, 1910) Thomson, 1858	68	BC	Gusarov 2003e
		scolopacina Casey, 1911	Atheta (Adota Casey, 1910) Thomson, 1858	124	CA	Gusarov 2003e
		scortea Casey, 1911	Atheta (Adota Casey, 1910) Thomson, 1858	124	CA	Gusarov 2003e
		setositarsis Casey, 1910	Atheta (Panalota Casey, 1910) Thomson, 1858	71	CA	Gusarov 2003e
		subintima Casey, 1910	Atheta (Adota Casey, 1910) Thomson, 1858	68	BC	Gusarov 2003e
Aloconota	brunneipes Casey,1906		Terasota Casey, 1906	337	NY
Aloconota	cambrica Wollaston, 1855		Atheta Thomson, 1858	205	Palaearctic Region	Klimaszewski & Winchester 2002
Aloconota	carmanah Klimaszewski in Klimaszewski & Winchester, 2002		Aloconota Thomson, 1858	20	BC	Klimaszewski & Winchester 2002
Aloconota	diversiseta Klimaszewski & Peck, 1986		Aloconota Thomson, 1858	64	AL	Klimaszewski & Peck 1986
Aloconota	flexibilis Casey, 1911		Aloconota Thomson, 1858	137	Mexico
Aloconota	neospelea Klimaszewski & Peck, 1986		Aloconota Thomson, 1858	65	AL	Klimaszewski & Peck 1986
Aloconota	perdita Casey, 1910		Aloconota (Terasota Casey, 1906 ) Thomson, 1858	85	CA
Aloconota	sulcifrons (Stephens, 1832)		Aleochara Gravenhorst, 1802	121	Palaearctic Region	Benick 1954
		insecta sensu Klimaszewsky & Peck, 1986	Aloconota Thomson, 1858	65		Gusarov 2003g
		laurentiana Blatchley, 1910	Atheta Thomson, 1858	357	IN	Gusarov 2003g
Aloconota	ventralis Casey, 1906		Taphrodota Casey, 1906	339	NY
Amischa	angusta Casey, 1893		Colposura Casey, 1893	339	NV
Amischa	colonia Casey, 1910		Amischa (s. str.) Thomson, 1858	98	CA
Amischa	continentalis Bernhauer, 1909		Atheta (Amischa Thomson, 1858) Thomson, 1858	528	IA
Amischa	devincta Casey, 1910		Amischa (s. str.) Thomson, 1858	98	CA
Amischa	morula Casey, 1911		Oxypoda Mannerheim, 1830	45	CA
Amischa	normalis Casey, 1910		Amischa (s. str.) Thomson, 1858	97	NY
Amischa	parviceps Casey, 1893		Colposura Casey, 1893	338	WA
Amischa	praelonga Casey, 1893		Colposura Casey, 1893	337	WY
Amischa	tersa Casey, 1910		Amischa (Colposura Casey, 1893) Thomson, 1858	99	CO
Anatheta	aspericauda Bernhauer, 1907a		Atheta (s.str.) Thomson, 1858	400	FL
Anatheta	georgiae Bernhauer et Scheerpeltz, 1926		Atheta (s.str.) Thomson, 1858	643	n. nov. for Atheta orientis Bernhauer, 1909 (nec Fauvel, 1904)
		orientis Bernhauer, 1909	Atheta (s.str.) Thomson, 1858	525	GA
Anatheta	planulicollis Casey, 1910		Sableta (Anatheta Casey, 1910) Casey, 1910	112	KS	Ashe & Gusarov 2003
		erudita Casey, 1911	Metaxya Mulsant & Rey, 1873	135	MB	Ashe & Gusarov 2003
Anatheta	surrufa Casey, 1911		Metaxya Mulsant & Rey, 1873	135	MB	Ashe & Gusarov 2003
Atheta	admista Casey, 1910		Aloconota (s. str.) Thomson, 1858	84	IA
Atheta	aemula Erichson, 1839		Homalota Mannerheim, 1830	102	PA	Gusarov 2003g
		bucolica Casey, 1910	Atheta (s.str.) Thomson, 1858	46	MS	Gusarov 2003g
		disjuncta Casey, 1910	Atheta (s.str.) Thomson, 1858	43	PA	Gusarov 2003g
		euryptera sensu Bernhauer, 1907a	Atheta (s.str.) Thomson, 1858	392	PA	Gusarov 2003g
		polita Melsheimer, 1844	Homalota Mannerheim, 1830	31	PA	Gusarov 2003g
		replicans Casey, 1910	Atheta (s.str.) Thomson, 1858	44	IA	Gusarov 2003g
		spadix Casey, 1910	Atheta (s.str.) Thomson, 1858	45	NY	Gusarov 2003g
Atheta	alabama Klimaszewski & Peck, 1986		Atheta (s.str.) Thomson, 1858	76	AL	Klimaszewski & Peck 1986
Atheta	alamedana Casey, 1910		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	59	CA
Atheta	altaica Bernhauer, 1901		Atheta Thomson, 1858	109	Palaearctic Region	Lohse et al. 1990
Atheta	amens Casey, 1911		Atheta (s.str.) Thomson, 1858	97	PA
Atheta	amicula Stephens, 1832		Aleochara Gravenhorst, 1802	132	Palaearctic Region	Benick & Lohse 1974
Atheta	annexa Casey, 1910		Atheta (s.str.) Thomson, 1858	43	NC	Klimaszewski & Peck 1986
		burra Casey, 1911	Atheta (s.str.) Thomson, 1858	100	WI	Gusarov 2003g
		citata Casey, 1910	Atheta (s.str.) Thomson, 1858	41	NY	Gusarov 2003g
		evecta Casey, 1910	Atheta (s.str.) Thomson, 1858	44	MS	Gusarov 2003g
		nacta Casey, 1911	Atheta (s.str.) Thomson, 1858	101	MO	Gusarov 2003g
		palpator Casey, 1911	Atheta (s.str.) Thomson, 1858	99	OH	Gusarov 2003g
		propitia Casey, 1911	Atheta (s.str.) Thomson, 1858	99	NC	Gusarov 2003g
Atheta	aperta Casey, 1910		Atheta (s.str.) Thomson, 1858	33	CA
		intacta Casey, 1910	Atheta (s.str.) Thomson, 1858	110	CA
		morbosa Casey, 1911	Atheta (s.str.) Thomson, 1858	107	CA
Atheta	astuta Casey, 1910		Atheta (Philhygra Mulsant & Rey, 1873) Thomson, 1858	65	PA
		nugatoria Casey, 1911	Pseudota Casey, 1910	149	PA
Atheta	atomica Casey, 1911		Datomicra (Hilarina Casey, 1910) Mulsant & Rey, 1874	153	NY
Atheta	audens Casey, 1911		Atheta (s.str.) Thomson, 1858	107	CA
		lepidula Casey, 1911	Atheta (s.str.) Thomson, 1858	108	CA
Atheta	bakeri Bernhauer, 1909		Atheta (Acrotona Thomson, 1859) Thomson, 1858	526	PA
Atheta	baringiana Bernhauer, 1907a		Atheta (Datomicra Mulsant & Rey, 1874) Thomson, 1858	396	WA
Atheta	bidenticulata Bernhauer, 1907a		Atheta (s.str.) Thomson, 1858	399	CA
Atheta	blatchleyi Bernhauer & Scheerpeltz, 1926		Atheta (s.str.) Thomson, 1858	639	n. nov. for Atheta caviceps Blatchley, 1910 (nec Poppius, 1909)	Lohse et al. 1990 (as Boreophilia chillcotti)
		caviceps Blatchley, 1910	Atheta Thomson, 1858	354	IN	Gusarov 2003g
		chillcotti Lohse in Lohse et al., 1990	Boreophilia Benick, 1973	157	MB	Gusarov 2003g
Atheta	brumalis Casey, 1910		Atheta (s.str.) Thomson, 1858	32	BC	Klimaszewski & Winchester 2002
		repexa Casey, 1911	Atheta (s.str.) Thomson, 1858	109	CA	Gusarov 2003g
Atheta	burwelli Lohse in Lohse et al., 1990		Dimetrota Mulsant & Rey, 1873	178	PQ	Lohse et al. 1990
Atheta	californica Bernhauer, 1907a		Atheta (Metaxya Mulsant & Rey, 1873) Thomson, 1858	385	CA
Atheta	campbelli Lohse in Lohse et al., 1990		Dimetrota Mulsant & Rey, 1873	185	YT	Lohse et al. 1990
Atheta	campbelliana Lohse in Lohse et al., 1990		Boreostiba Lohse in Lohse et al., 1990	195	YT	Lohse et al. 1990
Atheta	caribou Lohse in Lohse et al., 1990		Dimetrota Mulsant & Rey, 1873	182	YT	Lohse et al. 1990
Atheta	catula Casey, 1910		Atheta (s.str.) Thomson, 1858	94	NY
Atheta	centropunctata Bernhauer, 1909		Atheta (Metaxya Mulsant & Rey, 1873) Thomson, 1858	515	CA
Atheta	cheersae Klimaszewski in Klimaszewski & Winchester, 2002		Atheta (Metadimetrota Klimaszewski in Klimaszewski & Winchester, 2002) Thomson, 1858	32	BC	Klimaszewski & Winchester 2002
Atheta	circulicollis Lohse in Lohse et al., 1990		Atheta Thomson, 1858	189	PQ	Lohse et al. 1990
Atheta	claricella Casey, 1910		Atheta (s.str.) Thomson, 1858	31	CA
		dunni Casey, 1910	Atheta (s.str.) Thomson, 1858	31	CA
Atheta	clienta Casey, 1911		Pseudota Casey, 1910	150	MS
Atheta	concessa Casey, 1911		Atheta (Lamiota Casey, 1910) Thomson, 1858	83	BC	Klimaszewski & Winchester 2002
		sumpta Casey, 1911	Atheta (s.str.) Thomson, 1858	92	BC	Gusarov 2003g
Atheta	cornelli Pace, 1997		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	106	OR	Pace 1997
Atheta	coruscula Casey, 1910		Datomicra Mulsant & Rey, 1874	124	CA
Atheta	crenulata Bernhauer, 1907a		Atheta (s.str.) Thomson, 1858	391	WA
Atheta	crenuliventris Bernhauer, 1907a		Atheta (s.str.) Thomson, 1858	393	ME	Gusarov 2003f
		bradorensis Lohse in Lohse et al., 1990	Dimetrota Mulsant & Rey, 1873	179	PQ	Gusarov 2003f
Atheta	cryptica Lohse in Lohse et al., 1990		Bessobia Thomson, 1858	197	PQ
Atheta	cursor Mäklin in Mannerheim, 1852		Homalota Mannerheim, 1830	307	AK	Lohse & Smetana 1985
Atheta	dadopora Thomson, 1867		Atheta Thomson, 1858	283	Palaearctic Region	Benick & Lohse 1974
		decolorata Casey, 1910	Datomicra Mulsant & Rey, 1874	120	NY	Gusarov 2003g
		inopia Casey, 1910	Datomicra Mulsant & Rey, 1874	120	NY	Gusarov 2003g
		schematica Casey, 1910	Datomicra Mulsant & Rey, 1874	121	PA	Gusarov 2003g
		stilla Casey, 1910	Datomicra Mulsant & Rey, 1874	123	RI	Gusarov 2003g
Atheta	dama Casey, 1910		Atheta (s.str.) Thomson, 1858	27	NY
Atheta	delumbis Casey, 1911		Atheta (Delphota Casey, 1910) Thomson, 1858	87	NJ
Atheta	demissa Notman, 1921		Amphibitherion Notman, 1921	155	NY
Atheta	diffidens Casey, 1910		Datomicra Mulsant & Rey, 1874	122	NY
Atheta	discipula Casey, 1910		Atheta (s.str.) Thomson, 1858	33	NY
		jerseyiana Bernhauer & Scheerpeltz, 1926	Atheta (Philhygra Mulsant & Rey, 1873) Thomson, 1858	629	n. nov. for Atheta repanda Casey, 1910 (nec Mulsant & Rey, 1874)
		repanda Casey, 1910	Atheta (Philhygra Mulsant & Rey, 1873) Thomson, 1858	65	NJ
Atheta	disparilis Casey, 1910		Synaptina Casey, 1910	133	CA
Atheta	districta Casey, 1911		Atheta (s.str.) Thomson, 1858	113	BC
Atheta	enitescens Casey, 1910		Atheta (s.str.) Thomson, 1858	20	NY
Atheta	esmeraldae Casey, 1911		Atheta (s.str.) Thomson, 1858	116	NV
Atheta	fanatica Casey, 1910		Atheta (s.str.) Thomson, 1858	38	BC
		omissa Casey, 1910	Dimetrota Mulsant & Rey, 1873	101	BC
		vicaria Casey, 1910	Dimetrota Mulsant & Rey, 1873	101	BC
Atheta	fascinans Casey, 1911		Pseudota Casey, 1910	148	NY
Atheta	fenyesi Bernhauer, 1907a		Atheta (Dimetrota Mulsant & Rey, 1873) Thomson, 1858	395	CA	Gusarov 2003g
		cerebrosa Casey, 1911	Dimetrota (s.str.) Mulsant & Rey, 1873	142	CA	Gusarov 2003g
		immerita Casey, 1911	Dimetrota (s. str.) Mulsant & Rey, 1873)	141	CA	Gusarov 2003g
		incredula Casey, 1911	Atheta (Dimetrota Mulsant & Rey, 1873) Thomson, 1858	141	CA	Gusarov 2003g
		opinata Casey, 1911	Dimetrota (s.str.) Mulsant & Rey, 1873	142	CA	Gusarov 2003g
		querula Casey, 1910	Atheta (s.str.) Thomson, 1858	36	CA	Gusarov 2003g
		resima Casey, 1910	Dimetrota (s.str.) Mulsant & Rey, 1873	100	CA	Gusarov 2003g
		socors Casey, 1911	Atheta (s.str.) Thomson, 1858	108	CA	Gusarov 2003g
		vigilans Casey, 1910	Dimetrota (s.str.) Mulsant & Rey, 1873	103	CA	Gusarov 2003g
Atheta	festinans Erichson, 1839		Homalota Mannerheim, 1830	112	PA	Gusarov 2003g
		consonens Casey, 1910	Synaptina Casey, 1910	132	RI	Gusarov 2003g
		merica Casey, 1910	Synaptina Casey, 1910	131	IA	Gusarov 2003g
		punctata Blatchley, 1910	Atheta Thomson, 1858	355	IN	Gusarov 2003g
		tartarea Casey, 1910	Ousipalia des Gozis, 1886	168	RI
Atheta	ficta Casey, 1910		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	59	CA
Atheta	finita Moore & Legner, 1975		Atheta (Adota Casey, 1910) Thomson, 1858	361	n. nov. for Atheta definita Casey, 1911 (nec Casey, 1910)
		definita Casey, 1911	Atheta (Adota Casey, 1910) Thomson, 1858	123	CA
Atheta	flaviventris Casey, 1911		Sableta (Canastota Casey, 1910) Casey, 1910	111	TX
Atheta	formalis Casey, 1911		Pseudota Casey, 1910	152	BC
Atheta	freta Casey, 1910		Atheta (s.str.) Thomson, 1858	44	ND
Atheta	frosti Bernhauer, 1909		Atheta (s.str.) Thomson, 1858	520	MA	Gusarov 2003g
		candidula Casey, 1911	Atheta (s.str.) Thomson, 1858	94	NY	Gusarov 2003g
		comitata Casey, 1910	Atheta (s.str.) Thomson, 1858	24	RI	Gusarov 2003g
		diffisa Casey, 1911	Atheta (s.str.) Thomson, 1858	95	NY	Gusarov 2003g
		elota Casey, 1910	Atheta (s.str.) Thomson, 1858	26	NY	Gusarov 2003g
		gnoma Casey, 1910	Atheta (s.str.) Thomson, 1858	25	NY	Gusarov 2003g
		insidiosa Casey, 1910	Atheta (s.str.) Thomson, 1858	26	NY	Gusarov 2003g
		modiella Casey, 1911	Atheta (s.str.) Thomson, 1858	96	RI	Gusarov 2003g
		nata Casey, 1911	Atheta (s.str.) Thomson, 1858	95	RI	Gusarov 2003g
		puricula Casey, 1911	Pseudota Casey, 1910	148	NY	Gusarov 2003g
		vacillans Casey, 1911	Atheta (s.str.) Thomson, 1858	98	RI	Gusarov 2003g
Atheta	fulgens Casey, 1907		Atheta (s.str.) Thomson, 1858	392	WV
		lubricans Casey, 1910	Acrotona (Neada Casey, 1910) Thomson, 1859	153	RI
Atheta	fulgida Bernhauer, 1907a		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	399	CA
Atheta	fulviceps Notman, 1920		Atheta (Hilara Mulsant & Rey, 1873) Thomson, 1858	726	FL
Atheta	graminicola Gravenhorst, 1806		Aleochara Gravenhorst, 1802	176	Palaearctic Region	Benick & Lohse 1974
		granulata Mannerheim, 1846	Stonalota Mannerheim, 1846	508	AK	Gusarov 2003g
		lanei Casey, 1910	Atheta (Homalotusa Casey, 1906) Thomson, 1858	10	OR	Gusarov 2003g
		nomadica Casey, 1910	Atheta (Megista Mulsant & Rey, 1874) Thomson, 1858	15	AK	Gusarov 2003g
Atheta	hampshirensis Bernhauer, 1909		Atheta (Dimetrota Mulsant & Rey, 1873) Thomson, 1858	525	NH	Lohse & Smetana 1988; Klimaszewski & Winchester 2002
		hebescens Casey, 1910	Datomicra (s.str.) Mulsant & Rey, 1874	124	NY	Gusarov 2003g
		leviceps Casey, 1910	Atheta (Philhygra Mulsant & Rey, 1873) Thomson, 1858	66	NY	Gusarov 2003g
		maeklini Fenyes, 1920	Atheta (Dimetrota Mulsant & Rey, 1873) Thomson, 1858	205	n. nov. for Atheta moesta (Maeklin, 1852) (nec Zetterstedt, 1828)	Gusarov 2003g
		moesta Mäklin in Mannerheim, 1852	Homalota Mannerheim, 1830	307	AK	Gusarov 2003g
		pellax Casey, 1910	Datomicra (s.str.) Mulsant & Rey, 1874	124	RI	Gusarov 2003g
		retrusa Casey, 1910	Dimetrota (s.str.) Mulsant & Rey, 1873	103	BC	Gusarov 2003g
		sectator Casey, 1910	Dimetrota (s.str.) Mulsant & Rey, 1873	102	CA	Gusarov 2003g
		vana Casey, 1911	Pseudota Casey, 1910	150	BC	Gusarov 2003g
Atheta	hesperica Casey, 1910		Ousipalia des Gozis, 1886	169	CA
Atheta	hilaris Fenyes, 1909		Atheta (s.str.) Thomson, 1858	420	NM
Atheta	holmbergi Bernhauer, 1907a		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	389	AK	Klimaszewski & Winchester 2002
Atheta	houstoni Casey, 1910		Atheta (Philhygra Mulsant & Rey, 1873) Thomson, 1858	67	TX
Atheta	immunis Casey, 1910		Sableta (Taxicerella Casey, 1910) Casey, 1910	114	CA
Atheta	impressipennis Bernhauer, 1909		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	518	MA
Atheta	inanis Casey, 1910		Datomicra Mulsant & Rey, 1874	129	PA
Atheta	insolida Casey, 1910		Dimetrota Mulsant & Rey, 1873	125	CA
Atheta	intecta Casey, 1910		Aloconota (Terasota Casey, 1906) Thomson, 1858	86	NC
Atheta	invenusta Casey, 1910		Metaxya Mulsant & Rey, 1873	78	UT
Atheta	irrita Casey, 1911		Atheta (Adota Casey, 1910) Thomson, 1858	126	NV
Atheta	irrupta Casey, 1910		Pseudota Casey, 1910	117	BC	Klimaszewski & Winchester 2002
Atheta	kansana Casey, 1911		Atheta (s.str.) Thomson, 1858	92	KS
Atheta	keeni Casey, 1910		Atheta (Lamiota Casey, 1910) Thomson, 1858	17	BC	Klimaszewski & Winchester 2002 (as Atheta vasta)
		achromata Casey, 1911	Atheta (Lamiota Casey, 1910) Thomson, 1858	82	BC	Gusarov 2003g
		innocens Casey, 1910	Atheta (s.str.) Thomson, 1858	39	OR	Gusarov 2003g
		profecta Casey, 1911	Atheta (Lamiota Casey, 1910) Thomson, 1858	83	BC	Gusarov 2003g
Atheta	klagesi Bernhauer, 1909		Atheta (s.str.) Thomson, 1858	524	ME	Gusarov 2003g
		cephalina Casy, 1910	Atheta (Delphota Casey, 1910) Thomson, 1858	18	IA	Gusarov 2003g
		disca Moore & Legner, 1975	Atheta (s.str.) Thomson, 1858	359	n. nov. for Atheta discreta Casey, 1910: 42 (nec Casey, 1893, nec Casey, 1910: 79)	Gusarov 2003g
		discreta Casey, 1910	Atheta (s.str.) Thomson, 1858	42	NY	Gusarov 2003g
		dissensa Casey, 1910	Pseudota Casey, 1910	115	PA	Gusarov 2003g
		nympha Casey, 1910	Atheta (s.str.) Thomson, 1858	42	NY	Gusarov 2003g
		villica Casey, 1911	Atheta (s.str.) Thomson, 1858	97	PA	Gusarov 2003g
Atheta	laetula Fenyes, 1909		Atheta (Dimetrota Mulsant & Rey, 1873) Thomson, 1858	421	NM
Atheta	lagunae Lohse in Lohse et al., 1990		Boreostiba Lohse in Lohse et al., 1990	195	YT	Lohse et al. 1990
Atheta	limulina Casey, 1911		Atheta (s.str.) Thomson, 1858	93	RI
		rusticula Casey, 1911	Atheta (s.str.) Thomson, 1858	93	NY
Atheta	lippa Casey, 1911		Sipalia (Sonomota Casey, 1911) Mulsant & Rey, 1853	158	CA
Atheta	longiclava Casey, 1910		Sableta (Canastota Casey, 1910) Casey, 1910	110	RI
Atheta	longicornis Gravenhorst, 1802		Aleochara Gravenhorst, 1802	87	Palaearctic Region	Benick & Lohse 1974
Atheta	lucana Casey, 1910		Datomicra (Monadia Casey, 1910) Mulsant & Rey, 1874	130	CA
Atehta	lucifuga Klimaszewski & Peck, 1986		Atheta (Dimetrota Mulsant & Rey, 1873) Thomson, 1858	72	AL	Klimaszewski & Peck 1986
Atheta	luctifera Bernhauer, 1906		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	341	CA
Atheta	lymphatica Casey, 1911		Atheta (Delphota Casey, 1910) Thomson, 1858	84	IA
		concussa Casey, 1911	Atheta (s.str.) Thomson, 1858	116	WA
		subretracta Casey, 1911	Atheta (s.str.) Thomson, 1858	115	ID
Atheta	marcescens Casey, 1911		Atheta (Nemota Casey, 1910) Thomson, 1858	122	CO
Atheta	marinica Casey, 1910		Atheta (s.str.) Thomson, 1858	34	CA
Atheta	martini Lohse in Lohse et al., 1990		Atheta Thomson, 1858	189	YT	Lohse et al. 1990
Atheta	metlakatlana Bernhauer, 1909		Atheta (s.str.) Thomson, 1858	522	BC	Klimaszewski & Winchester 2002
Atheta	militaris Bernhauer, 1909		Atheta (Ceritaxa Mulsant & Rey, 1873) Thomson, 1858	519	CA
Atheta	mina Casey, 1910		Datomicra (Hilarina Casey, 1910) Mulsant & Rey, 1874	153	CT
Atheta	modesta Melsheimer, 1844		Homalota Mannerheim, 1830	31	PA	Gusarov 2003g
		auguralis Casey, 1911	Atheta (s.str.) Thomson, 1858	104	NY	Gusarov 2003g
		bifaria Casey, 1911	Atheta (s.str.) Thomson, 1858	104	DC	Gusarov 2003g
		capella Casey, 1910	Atheta (s.str.) Thomson, 1858	24	RI	Gusarov 2003g
		ducens Casey, 1910	Atheta (s.str.) Thomson, 1858	25	NY	Gusarov 2003g
		fenisex Casey, 1911	Atheta (s.str.) Thomson, 1858	102	RI	Gusarov 2003g
		logica Casey, 1911	Atheta (Delphota Casey, 1910) Thomson, 1858	86	PA	Gusarov 2003g
		rhodeana Casey, 1910	Atheta (s.str.) Thomson, 1858	24	RI	Gusarov 2003g
		temperans Casey, 1910	Atheta (s.str.) Thomson, 1858	27	DC	Gusarov 2003g
		tradita Casey, 1911	Atheta (s.str.) Thomson, 1858	101	NY	Gusarov 2003g
		vierecki Casey, 1911	Atheta (s.str.) Thomson, 1858	103	CT	Gusarov 2003g
		virginica Bernhauer, 1907a	Atheta (s.str.) Thomson, 1858	393	WV	Gusarov 2003g
Atheta	muensteri Bernhauer, 1902		Atheta Thomson, 1858	55	Palaearctic Region	Lohse et al. 1990
Atheta	nacta Casey, 1911		Atheta (s.str.) Thomson, 1858	101	MO
		burra Casey, 1911	Atheta (s.str.) Thomson, 1858	100	WI
		palpator Casey, 1911	Atheta (s.str.) Thomson, 1858	99	OH
Atheta	nanella Casey, 1906		Silusida Casey, 1906	271	PA
		beatula Casey, 1910	Sableta (Canastota Casey, 1910) Casey, 1910	109	PA	Ashe & Gusarov in press
		ornator Casey, 1910	Sableta (Canastota Casey, 1910) Casey, 1910	110	NY	Ashe & Gusarov in press
Atheta	nearctica Lohse in Lohse et al., 1990		Dimetrota Mulsant & Rey, 1873	187	YT	Lohse et al. 1990
Atheta	neomexicana Fenyes, 1909		Atheta (Dimetrota Mulsant & Rey, 1873) Thomson, 1858	422	NM
Atheta	nescia Casey, 1910		Pseudota Casey, 1910	117	BC
Atheta	nigrita Fenyes, 1909		Atheta (Dimetrota Mulsant & Rey, 1873) Thomson, 1858	423	CA
		incumbens Casey, 1910	Datomicra Mulsant & Rey, 1874	123	NV
		perpaula Casey, 1910	Datomicra (Oligomia Casey, 1910) Mulsant & Rey, 1874	130	CA
		pomonae Casey, 1910	Datomicra Mulsant & Rey, 1874	121	CA
		scintilla Casey, 1910	Datomicra (Oligomia Casey, 1910) Mulsant & Rey, 1874	129	CA
Atheta	nugator Casey, 1911		Atheta (Hilara Mulsant & Rey, 1873) Thomson, 1858	63	RI
Atheta	obsequens Casey, 1911		Atheta (Traumoecia Mulsant & Rey, 1874) Thomson, 1858	119	NY
Atheta	occidentalis Bernhauer, 1906		Atheta (s.str.) Thomson, 1858	338	CA
		civica Casey, 1910	Atheta (s.str.) Thomson, 1858	29	CA
		fugitans Casey, 1911	Atheta (s.str.) Thomson, 1858	106	CA
		prolata Casey, 1910	Atheta (s.str.) Thomson, 1858	37	CA
Atheta	ordinata Casey, 1910		Atheta (s.str.) Thomson, 1858	28	NY
Atheta	oregonensis Bernhauer, 1909		Atheta (s.str.) Thomson, 1858	523	OR
Atheta	pacifica Casey, 1910		Ousipalia des Gozis, 1886	169	CA
Atheta	paganella Casey, 1910		Atheta (Nemota Casey, 1910) Thomson, 1858	56	CA
		filiola Casey, 1910	Atheta (Nemota Casey, 1910) Thomson, 1858	57	CA
		informalis Casey, 1911	Atheta (Nemota Casey, 1910) Thomson, 1858	121	CA
		timida Casey, 1910	Atheta (Nemota Casey, 1910) Thomson, 1858	57	CA
Atheta	particula Casey, 1910		Datomicra (Hilarina Casey, 1910) Mulsant & Rey, 1874	128	RI
Atheta	parvipennis Bernhauer, 1907a		Atheta (Ousipalia des Gozis, 1886) Thomson, 1858	398	AB	Lohse et al. 1990 (as Boreostiba hudsonica)
		hudsonica Lohse in Lohse et al., 1990	Boreostiba Lohse in Lohse et al., 1990	193	PQ	Gusarov 2003g
Atheta	pavidula Casey, 1911		Atheta (Adota Casey, 1910) Thomson, 1858	126	CA
Atheta	pedicularis Melsheimer, 1944		Oligota Mannerheim, 1830	31	PA
Atheta	pennsylvanica Bernhauer, 1907a		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	388	MA	Gusarov 2003g
		fontis Casey, 1911	Atheta (Hilara Mulsant & Rey, 1873) Thomson, 1858	122	PA	Gusarov 2003g
Atheta	perpera Casey, 1910		Atheta (Nemota Casey, 1910) Thomson, 1858	58	CA
		meticulosa Casey, 1911	Atheta (Nemota Casey, 1910) Thomson, 1858	121	CA
		perspecta Casey, 1910	Atheta (Nemota Casey, 1910) Thomson, 1858	58	CA
Atheta	personata Casey, 1910		Noverota Casey, 1910	92	VA
Atheta	perversa Casey, 1910		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	61	CA
Atheta	picipennis Mannerheim, 1843		Homalota Mannerheim, 1830	224	AK	Klimaszewski & Winchester 2002
		alaskana Casey, 1911	Atheta (s.str.) Thomson, 1858	113	AK	Gusarov 2003g
		aperta Casey, 1910	Atheta (s.str.) Thomson, 1858	33	CA	Gusarov 2003g
		carlottae Casey, 1910	Atheta (s.str.) Thomson, 1858	32	BC	Gusarov 2003g
		intacta Casey, 1911	Atheta (s.str.) Thomson, 1858	110	CA	Gusarov 2003g
		morbosa Casey, 1911	Atheta (s.str.) Thomson, 1858	107	CA	Gusarov 2003g
		wrangelica Casey, 1911	Atheta (s.str.) Thomson, 1858	91	AK	Gusarov 2003g
Atheta	pimalis Casey, 1910		Pseudota Casey, 1910	118	AZ
Atheta	praesaga Casey, 1910		Pseudota Casey, 1910	116	CA
Atheta	pratensis Mäklin in Mannerheim, 1852		Homalota Mannerheim, 1830	308	AK	Lohse & Smetana 1985
Atheta	promota Casey, 1910		Atheta (s.str.) Thomson, 1858	36	MS
Atheta	properans Casey, 1910		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	60	CA
Atheta	prudhoensis Lohse in Lohse et al., 1990		Dimetrota Mulsant & Rey, 1873	182	AK	Lohse et al. 1990
Atheta		dempsterensis Lohse in Lohse et al., 1990	Dimetrota Mulsant & Rey, 1873	181	YT	Gusarov 2003g
Atheta	pseudoatomaria Bernhauer, 1909		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	518	ME
Atheta	pseudovilis Bernhauer, 1907a		Atheta (Dralica Mulsant & Rey, 1874) Thomson, 1858	386	CA
Atheta	pugnans Fenyes, 1920		Atheta (Traumoecia Mulsant & Rey, 1874) Thomson, 1858	218	n. nov. for Atheta militaris Casey, 1911 (nec Bernhauer, 1909)
		militaris Casey, 1911	Atheta (Traumoecia Mulsant & Rey, 1874) Thomson, 1858	118	AK
Atheta	quaesita Casey, 1910		Synaptina Casey, 1910	132	IA
Atheta	reformata Casey, 1911		Atheta (s.str.) Thomson, 1858	102	WI
Atheta	regenerans Casey, 1911		Atheta (Delphota Casey, 1910) Thomson, 1858	85	IA
Atheta	regissalmonis Lohse in Lohse et al., 1990		Dimetrota Mulsant & Rey, 1873	179	AK	Lohse et al. 1990
Atheta	relicta Casey, 1911		Atheta (s.str.) Thomson, 1858	112	BC
Atheta	remissa Casey, 1910		Sableta (Taxicerella Casey, 1910) Casey, 1910	113	PA
Atheta	remulsa Casey, 1910		Atheta (s.str.) Thomson, 1858	30	BC
		asperiola Casey, 1910	Atheta (Traumoecia Mulsant & Rey, 1874) Thomson, 1858	47	NY
		frugalis Casey, 1910	Atheta (Philhygra Mulsant & Rey, 1873) Thomson, 1858	66	NY
Atheta	reposita Casey, 1910		Atheta (s.str.) Thomson, 1858	35	UT
		arizonica Casey, 1910	Atheta (s.str.) Thomson, 1858	22	AZ
		ostenta Casey, 1910	Atheta (s.str.) Thomson, 1858	35	AZ
Atheta	restricta Casey, 1911		Atheta (Donesia Casey, 1910) Thomson, 1858	120	VA
		necessita Moore & Legner, 1975	Atheta (Donesia Casey, 1910) Thomson, 1858	367	n. nov. for Atheta restricta Casey, 1911 (nec Casey, 1910)
Atheta	reticula Casey, 1910		Atheta (s.str.) Thomson, 1858	40	VA
Atheta	ringi Klimaszewski in Klimaszewski & Winchester, 2002		Atheta (s.str.) Thomson, 1858	24	BC	Klimaszewski & Winchester 2002
Atheta	rurigena Casey, 1911		Atheta (s.str.) Thomson, 1858	114	BC
Atheta	sana Casey, 1910		Atheta (Micrearota Casey, 1910) Thomson, 1858	52	PA
Atheta	sitiens Casey, 1910		Pseudota Casey, 1910	116	IA
Atheta	smetanai Lohse in Lohse et al., 1990		Dimetrota Mulsant & Rey, 1873	183	PQ	Lohse et al. 1990
Atheta	sparreschneideri Munster, 1922		Atheta (s.str.) Thomson, 1858	207	Palaearctic Region	Lohse et al. 1990
Atheta	sparsepunctata Bernhauer, 1907a		Atheta (Ousipalia des Gozis, 1886) Thomson, 1858	398	CA
Atheta	stercoris Fenyes, 1920		Atheta (Dimetrota Mulsant & Rey, 1873) Thomson, 1858	205	n. nov. for Atheta columbica Bernhauer, 1907a (nec Fauvel, 1901)
		columbica Bernhauer, 1907a	Atheta (Dimetrota Mulsant & Rey, 1873) Thomson, 1858	394	BC
Atheta	stoica Casey, 1911		Atheta (s.str.) Thomson, 1858	110	CA
Atheta	strigosula Casey, 1910		Atheta (s.str.) Thomson, 1858	30	NY
Atheta	sublucens Bernhauer, 1909		Atheta (s.str.) Thomson, 1858	522	CA
		loquax Casey, 1911	Atheta (s.str.) Thomson, 1858	117	CA
		neutralis Casey, 1910	Atheta (s.str.) Thomson, 1858	38	CA
		vacunalis Casey, 1910	Dimetrota (s. str.) Mulsant & Rey, 1873	102	CA
Atheta	surgens Casey, 1910		Datomicra Mulsant & Rey, 1874	125	BC
Atheta	temporalis Casey, 1910		Atheta (Donesia Casey, 1910) Thomson, 1858	48	RI
Atehta	tepida Casey, 1910		Iotota Casey, 1910	95	CA
Atheta	texana Casey, 1910		Atheta (Stethusa Casey, 1910) Thomson, 1858	6	TX	Gusarov 2003f
		mendosa Casey, 1910	Atheta (Stethusa Casey, 1910) Thomson, 1858	8	VA	Gusarov 2003f
Atheta	tractabilis Casey, 1910		Atheta (s.str.) Thomson, 1858	27	NY
Atheta	troglophila Klimaszevsky & Peck, 1986		Atheta (Dimetrota Mulsant & Rey, 1873) Thomson, 1858	69	TN	Klimaszewski & Peck 1986
Atheta	truncativentris Bernhauer, 1907a		Atheta (s.str.) Thomson, 1858	400	CA
Atheta	tubericauda Bernhauer, 1909		Atheta (s.str.) Thomson, 1858	520	MA
		nexa Casey, 1910	Atheta (s.str.) Thomson, 1858	29	PA
Atheta	turpicula Cassey, 1910		Ousipalia des Gozis, 1886	170	CO
Atheta	umbonalis Casey, 1910		Atheta (s.str.) Thomson, 1858	22	OH
Atheta	unica Casey, 1910		Iotota Casey, 1910	95	CA
		sierranevadensis Likovsky, 1984	Atheta (Iotota Casey, 1910) Thomson, 1858	7	n. nov. for Atheta unica (Casey, 1910) (nec Bernhauer, 1907b)
Atheta	unigena Casey, 1910		Atheta (Hilara Mulsant & Rey, 1873) Thomson, 1858	62	NY
Atheta	vacans Casey, 1910		Datomicra Mulsant & Rey, 1874	122	NY
		filiformis Casey, 1910	Datomicra (Micromota Casey, 1910) Mulsant & Rey, 1874	127	NY
Atheta	vancouveri Klimaszewski in Klimaszewski & Winchester, 2002		Atheta (Datomicra Mulsant & Rey, 1874) Thomson, 1858	23	BC	Klimaszewski & Winchester 2002
Atheta	ventricosa Bernhauer, 1907a		Atheta (s.str.) Thomson, 1858	391	NJ	Gusarov 2003g
		callens Casey, 1911	Atheta (Delphota Casey, 1910) Thomson, 1858	87	DC	Gusarov 2003g
		franklini Casey, 1911	Atheta (s.str.) Thomson, 1858	89	PA	Gusarov 2003g
		mollicula Casey, 1910	Atheta (s.str.) Thomson, 1858	37	BC	Gusarov 2003g
		phrenetica Casey, 1910	Sableta (Canastota Casey, 1910) Casey, 1910	111	BC	Gusarov 2003g
		postulans Casey, 1911	Atheta (s.str.) Thomson, 1858	90	BC	Gusarov 2003g
Atheta	weedi Casey, 1910		Atheta (s.str.) Thomson, 1858	21	MS
Atheta	wrangeli Casey, 1910		Datomicra (Micromota Casey, 1910) Mulsant & Rey, 1874	126	AK
		nanulina Casey, 1911	Pseudota Casey, 1910	152	BC
Boreophilia	angusticornis Bernhauer, 1907a		Atheta (Metaxya Mulsant & Rey, 1873) Thomson, 1858	384	NH	Gusarov 2003g
		plutonica Casey, 1910	Metaxya Mulsant & Rey, 1873	82	NH	Gusarov 2003g
Boreophilia	caseyi Lohse in Lohse et al.E441		Boreophilia Benick, 1973	155	AK	Lohse et al. 1990
Boreophilia	eremita Rye, 1866		Homalota Mannerheim, 1830	123	Palaearctic Region	Lohse et al. 1990
Boreophilia	fusca C.R.Sahlberg, 1831		Aleochara Gravenhorst, 1802	371	Palaearctic Region	Lohse et al. 1990
Boreophilia	gelida J.Sahlberg, 1887		Atheta Thomson, 1858	26	Palaearctic Region	Lohse et al. 1990
Boreophilia	hyperborea Brundin, 1940a		Atheta Thomson, 1858	131	Palaearctic Region	Brundin 1940a
Boreophilia	insecuta Eppelsheim, 1893		Atheta Thomson, 1858	31	Palaearctic Region	Lohse et al. 1990
Boreophilia	islandica Kraatz, 1857		Homalota Mannerheim, 1830	284	Palaearctic Region	Lohse et al. 1990
Boreophilia	manitobensis Lohse in Lohse et al., 1990		Boreophilia Benick, 1973	160	MB	Lohse et al. 1990
Boreophilia	nearctica Lohse in Lohse et al., 1990		Boreophilia Benick, 1973	153	YT	Lohse et al. 1990
Boreophilia	nomensis Casey, 1910		Dinaraea Thomson, 1858	96	AK	Lohse et al. 1990 (as Boreophilia caseyiana)
		caseyiana Lohse in Lohse et al., 1990	Boreophilia Benick, 1973	160	YT	Gusarov 2003g
Boreophilia	subplana J.Sahlberg, 1880		Atheta Thomson, 1858	90	Palaearctic Region	Lohse et al. 1990
Boreophilia	venti Lohse in Lohse et al., 1990		Dimetrota Mulsant & Rey, 1873	183	YT	Lohse et al. 1990
Boreostiba	frigida J.Sahlberg, 1880		Atheta Thomson, 1858	93	Palaearctic Region	Brundin 1940b
		lamellifera Lohse in Lohse et al., 1990	Boreostiba Lohse in Lohse et al., 1990	195	PQ	Gusarov 2003g
Boreostiba	sibirica Mäklin, 1880		Homalota Mannerheim, 1830	82	Palaearctic Region	Brundin 1940b
Callicerus	obscurus Gravenhorst, 1802		Callicerus Gravenhorst, 1802	66	Palaearctic Region	Benick & Lohse 1974
Charoxus	spinifer Frank in Frank & Thomas, 1996		Charoxus Sharp, 1883	71	FL
Clusiota	impressicollis Bernhauer, 1907a		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	389	WA
		claviventris Casey, 1910	Clusiota Casey, 1910	119	BC
		nova Fenyes, 1920	Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	187	n. nov. for Atheta impressicollis (Bernhauer, 1907a) (nec Mulsant & Rey, 1852)
Crephalia	recessa Casey, 1910		Atheta (Crephalia Casey, 1910) Thomson, 1858	55	NY
		lineatula Casey, 1910	Sipalia Mulsant & Rey, 1853	167	IA
Dalotia	coriaria Kraatz, 1858		Homalota Mannerheim, 1830	282	Palaearctic Region	Benick & Lohse 1974
		crucialis Casey, 1910	Dimetrota (Dalotia Casey, 1910) Mulsant & Rey, 1873	107	CA	Gusarov 2003g
		miscella Casey, 1910	Pseudota Casey, 1910	115	NJ	Gusarov 2003g
		pectorina Casey, 1910	Dimetrota (Dalotia Casey, 1910) Mulsant & Rey, 1873	106	CA	Gusarov 2003g
Dilacra	cauta Casey, 1911		Oxypoda Mannerheim, 1830	23	CO
Dilacra	lineata Casey, 1893		Oxypoda Mannerheim, 1830	297	RI
Dilacra	vieta Casey, 1910		Omegalia Casey, 1910	94	CA
		abjecta Casey, 1910	Omegalia Casey, 1910	94	CA
Dinaraea	angustula Gyllenhal, 1810		Aleochara Gravenhorst, 1802	393	Palaearctic Region	Benick & Lohse 1974
Dinaraea	borealis Lohse in Lohse et al., 1990		Dinaraea Thomson, 1858	198	PQ	Lohse et al. 1990
Dinaraea	planaris Mäklin in Mannerheim, 1852		Homalota Mannerheim, 1830	309	AK	Lohse & Smetana 1985
Dinaraea	subdepressa Bernhauer, 1907a		Atheta (Dinaraea Thomson, 1858) Thomson, 1858	386	NH
Dochmonota	rudiventris Eppelsheim, 1886		Homalota Mannerheim, 1830	35	Palaearctic Region	Benick & Lohse 1974 (as Atheta rudiventris)
		revoluta Casey, 1910	Dimetrota (s. str.) Mulsant & Rey, 1873	104	ID	Gusarov 2003g
		vaciva Casey, 1910	Datomicra (s. str.) Mulsant & Rey, 1874	126	MA	Gusarov 2003g
Earota	dentata Bernhauer, 1906		Atheta Thomson, 1858	342	CA	Gusarov 2002d
		alutacea Casey, 1906	Macroterma Casey, 1906	335	NY	Gusarov 2002d
		borealis Casey, 1906	Macroterma Casey, 1906	336	NH	Gusarov 2002d
		iowensis Casey, 1910	Atheta (Macroterma Casey, 1906) Thomson, 1858	10	IA	Gusarov 2002d
Emmelostiba	microptera Lohse in Lohse et al., 1990		Pseudousipalia Lohse in Lohse et al., 1990	150	YT	Lohse et al. 1990
Euromota	lucida Casey, 1906		Euromota Casey, 1906	338	VA
		euromota Likovsky, 1984	Atheta (Euromota Casey, 1906) Thomson, 1858	7	n. nov. for Atheta lucida (Casey, 1906) (nec Dodero, 1922)
Gaenima	impedita Casey, 1911		Gaenima Casey, 1911	161	CA
Geostiba (Sibiota)	alticola Lohse & Smetana, 1988		Geostiba Thomson, 1858	271	NC	Gusarov 2002c
Geostiba (Sibiota)	appalachigena Gusarov, 2002c		Geostiba (Sibiota Casey, 1906) Thomson, 1858	19	WV	Gusarov 2002c
Geostiba (Sibiota)	balsamensis Pace, 1997		Geostiba (Lioglutosipalia Scheerpeltz, 1951) Thomson, 1858	104	NC	Gusarov 2002c
Geostiba (Sibiota)	bicarinata Lohse & Smetana, 1988		Geostiba Thomson, 1858	273	NC	Gusarov 2002c
Geostiba (Sibiota)	carteriensis Pace, 1997		Geostiba (Lioglutosipalia Scheerpeltz, 1951) Thomson, 1858	106	NC/TN	Gusarov 2002c
Geostiba (s. str.)	circellaris Gravenhorst, 1806		Aleochara Gravenhorst, 1802	155	Palaearctic Region	Gusarov 2002c
Geostiba (Sibiota)	coeligena Gusarov, 2002c		Geostiba (Sibiota Casey, 1906) Thomson, 1858	57	NC/TN	Gusarov 2002c
Geostiba (Sibiota)	crepusculigena Gusarov, 2002c		Geostiba (Sibiota Casey, 1906) Thomson, 1858	44	TN	Gusarov 2002c
Geostiba (Sibiota)	graveyardensis Pace, 1997		Geostiba (Ditroposipalia Scheerpeltz, 1951) Thomson, 1858	102	NC	Gusarov 2002c
Geostiba (Sibiota)	impressula Casey, 1906		Sibiota Casey, 1906	350	OR	Gusarov 2002c
		fossata Casey, 1910	Sipalia Mulsant & Rey, 1853	167	OR	Gusarov 2002c
Geostiba (Sibiota)	nebuligena Gusarov, 2002c		Geostiba (Sibiota Casey, 1906) Thomson, 1858	40	NC/TN	Gusarov 2002c
Geostiba (Sibiota)	nimbicola Lohse & Smetana, 1988		Geostiba Thomson, 1858	275	NC/TN	Gusarov 2002c
Geostiba (Sibiota)	nubigena Lohse & Smetana, 1988		Geostiba Thomson, 1858	273	NC	Gusarov 2002c
Geostiba (Sibiota)	pluvigena Gusarov, 2002c		Geostiba (Sibiota Casey, 1906) Thomson, 1858	36	NC	Gusarov 2002c
Geostiba (Sibiota)	silvigena Gusarov, 2002c		Geostiba (Sibiota Casey, 1906) Thomson, 1858	72	CA	Gusarov 2002c
Goniusa	alperti Kistner, 1976		Goniusa Casey, 1906	89	WA	Gusarov 2003b
Goniusa	caseyi Gusarov, 2003b		Goniusa Casey, 1906	9	MA	Gusarov 2003b
Halobrecta	algophila Fenyes, 1909		Atheta (Halobrectha Thomson, 1858) Thomson, 1858	419	CA	Gusarov in press, a
		barbarae Casey, 1910	Atheta (s.str.) Thomson, 1858	18	CA	Gusarov in press, a
		importuna Casey, 1911	Atheta (s.str.) Thomson, 1858	111	CA	Gusarov in press, a
Halobrecta	flavipes Thomson, 1861		Halobrecta Thomson, 1858	50	Palaearctic Region	Gusarov in press, a
		incertula Casey, 1910	Aloconota (s.str.) Thomson, 1858	84	VA	Gusarov in press, a
		pocahontas Casey, 1910	Atheta (s.str.) Thomson, 1858	19	VA	Gusarov in press, a
		vaticina Casey, 1910	Atheta (s.str.) Thomson, 1858	19	NY	Gusarov in press, a
Halobrecthina	opaciceps Bernhauer, 1909		Atheta (Halobrecthina Bernhauer, 1909) Thomson, 1858	519	MA
		cribraticeps Casey, 1910	Atheta (Rovalida Casey, 1910) Thomson, 1858	69	RI
		monroei Casey, 1910	Atheta (Rovalida Casey, 1910) Thomson, 1858	71	VA
		piceicornis Casey, 1910	Atheta (Rovalida Casey, 1910) Thomson, 1858	70	NJ
		submaritima Casey, 1910	Atheta (Rovalida Casey, 1910) Thomson, 1858	70	NJ
Halobrecthina	sibylla Casey, 1910		Atheta (s.str.) Thomson, 1858	21	VA
Homia	gigantula LeConte, 1877		Thinobius Kiesenwetter, 1844	239	TX
Hydrosmecta	caduca Casey, 1910		Hydrosmecta Thomson, 1858	87	NY
Hydrosmecta	callidula Casey, 1910		Hydrosmecta Thomson, 1858	87	CA
		veris Fenyes, 1921	Lipodonta Fenyes, 1921	24	CA
Hydrosmecta	delicata Bernhauer, 1906		Atheta (Parameotica Ganglbauer, 1895) Thomson, 1858	340	CA
Hydrosmecta	depresseola Casey, 1911		Hydrosmecta Thomson, 1858	140	Mexico
Hydrosmecta	dulcis Casey, 1910		Hydrosmecta Thomson, 1858	87	WI
Hydrosmecta	fastidiosa Casey, 1911		Hydrosmecta Thomson, 1858	139	CA
Hydrosmecta	macra Fenyes, 1921		Hydrosmectina Ganglbauer, 1895	23	CA
Hydrosmecta	odiosica Casey, 1911		Hydrosmecta Thomson, 1858	138	CA
Hydrosmecta	pseudodiosica Lohse in Lohse et al., 1990		Hydrosmecta Thomson, 1858	149	YT	Lohse et al. 1990
Hydrosmecta	salinasica Casey, 1911		Hydrosmecta Thomson, 1858	139	CA
Hydrosmecta	subparilis Casey, 1910		Hydrosmecta Thomson, 1858	89	CA
Hydrosmecta	tincta Notman, 1921		Hydrosmecta Thomson, 1858	156	NY
		tinctina Moore & Legner, 1975	Atheta (Hydrosmecta Thomson, 1858) Thomson, 1858	375	n. nov. for Atheta tincta (Notman, 1921) (nec Casey, 1910)
Hydrosmecta	torrida Notman, 1921		Hydrosmecta Thomson, 1858	156	NY
Liogluta	abdominalis Bernhauer, 1907a		Atheta (Liogluta Thomson, 1858) Thomson, 1858	394	CA
Liogluta	aloconotoides Lohse in Lohse et al., 1990		Liogluta (Anepsiota Casey, 1893) Thomson, 1858	165	LB	Lohse et al. 1990
Liogluta	atriventris Casey, 1906		Athetota Casey, 1906	337	BC
Liogluta	blandita Casey, 1910		Atheta (s.str.) Thomson, 1858	32	CA
Liogluta	granulosa Lohse in Lohse et al., 1990		Liogluta (s. str.) Thomson, 1858	164	AK	Lohse et al. 1990
Liogluta	insignis Casey, 1885		Oxypoda Mannerheim, 1830	310	CA
		caseyi Bernhauer & Scheerpeltz, 1926	Atheta (Liogluta Thomson, 1858) Thomson, 1858	656	n. nov. for Atheta insignis (Casey, 1885) (nec Wollaston, 1854)
Liogluta	nevadensis Casey, 1910		Atheta (s.str.) Thomson, 1858	23	NV
Liogluta	nigropolita Bernhauer, 1907a		Atheta (Pseudomegista Bernhauer, 1907a) Thomson, 1858	390	NH	Lohse et al. 1990
Liogluta	nitens Mäklin in Mannerheim, 1852		Homalota Mannerheim, 1830	307	AK	Lohse & Smetana 1985, Klimaszewski & Winchester 2002
		apposita Casey, 1911	Atheta (s.str.) Thomson, 1858	90	BC	Gusarov 2003g
		insolens Casey, 1910	Atheta (Liogluta Thomson, 1858) Thomson, 1858	16	BC	Gusarov 2003g
		resplendens Casey, 1910	Dimetrota Mulsant & Rey, 1873	104	BC	Gusarov 2003g
Liogluta	novicia Casey, 1911		Atheta (s.str.) Thomson, 1858	105	CA
Liogluta	quadricollis Casey, 1893		Anepsiota Casey, 1893	330	BC
Liogluta	shastana Casey, 1910		Atheta (Anepsiota Casey, 1893) Thomson, 1858	13	CA
Liogluta	terminalis Casey, 1906		Anepsiota Casey, 1893	340	BC
		renominata Bernhauer & Scheerpeltz, 1926	Atheta (Liogluta Thomson, 1858) Thomson, 1858	658	n. nov. for Atheta terminalis (Casey, 1885) (nec Gravenhorst, 1806)
Liogluta	torpens Casey, 1910		Atheta (Anepsiota Casey, 1893) Thomson, 1858	13	CA
Liogluta	trapezicollis Lohse in Lohse et al., 1990		Liogluta (Anepsiota Casey, 1893) Thomson, 1858	165	YT	Lohse et al. 1990
Liogluta	vasta Maeklin, 1853		Homalota Mannerheim, 1830	183	AK	Lohse & Smetana 1985
Liogluta	wickhami Casey, 1893		Anepsiota Casey, 1893	331	BC
Lypoglossa	angularis angularis Mäklin in Mannerheim, 1853		Myrmedonia Erichson, 1837	181	AK	Gusarov in press, b
Lypoglossa	angularis fenyesi Bernhauer, 1907a		Dasyglossa Kraatz, 1856	402	BC	Gusarov in press, b
Lypoglossa	angularis obtusa LeConte, 1866		Goniusa Casey, 1906	373	PA	Gusarov in press, b
Lypoglossa	franclemonti Hoebeke, 1992		Lypoglossa Fenyes, 1918	386	NY	Gusarov in press, b
Lypoglossa	manitobae Gusarov, 2003		Lypoglossa Fenyes, 1918		MB	Gusarov in press, b
Micratheta	caudex Casey, 1910		Atheta (Micratheta Casey, 1910) Thomson, 1858	53	VA
Micrearota	loricula Casey, 1910		Atheta (Micrearota Casey, 1910) Thomson, 1858	49	IA
Micrearota	prolongata Casey, 1910		Atheta (Crephalia Casey, 1910) Thomson, 1858	55	MO
Micrearota	testata Casey, 1910		Atheta (Crephalia Casey, 1910) Thomson, 1858	56	PA
Micrearota	vestigialis Erichson, 1839		Homalota Mannerheim, 1830	112	PA
		definita Casey, 1910	Atheta (Micrearota Casey, 1910) Thomson, 1858	50	TX
		fecunda Casey, 1910	Atheta (Micrearota Casey, 1910) Thomson, 1858	52	NY
		gregaria Casey, 1910	Atheta (Micrearota Casey, 1910) Thomson, 1858	51	IA
		novagregaria Moore & Legner, 1975	Atheta (Stethusa Casey, 1910) Thomson, 1858	362	n. nov. for Atheta gregaria Casey, 1910 (nec Erichson, 1839)
		pristina Casey, 1910	Atheta (Micrearota Casey, 1910) Thomson, 1858	51	RI
		reperta Casey, 1910	Atheta (Micrearota Casey, 1910) Thomson, 1858	50	IA
		tincta Casey, 1910	Atheta (Micrearota Casey, 1910) Thomson, 1858	53	NY
		versuta Casey, 1910	Atheta (Micrearota Casey, 1910) Thomson, 1858	50	RI
Mocyta	amblystegii Brundin, 1952		Atheta Thomson, 1858	135	Palaearctic Region	Brundin 1952
Mocyta	breviuscula Maeklin in Mannerheim, 1852		Homalota Mannerheim, 1830	309	AK	Lohse & Smetana 1985
		ardelio Casey, 1910	Acrotona (s. str.) Thomson, 1859	150	CA	Gusarov 2003g
		digesta Casey, 1910	Acrotona (s. str.) Thomson, 1859	148	CA	Gusarov 2003g
		malaca Casey, 1910	Acrotona (s. str.) Thomson, 1859	151	CA	Gusarov 2003g
		prudens Casey, 1910	Acrotona (s. str.) Thomson, 1859	149	BC	Gusarov 2003g
		renoica Casey, 1910	Acrotona (s. str.) Thomson, 1859	150	NV	Gusarov 2003g
		severa Casey, 1910	Acrotona (s. str.) Thomson, 1859	148	CA	Gusarov 2003g
		shastanica Casey, 1910	Acrotona (s. str.) Thomson, 1859	148	CA	Gusarov 2003g
Mocyta	fungi Gravenhorst, 1806		Aleochara Gravenhorst, 1802	157	Palaearctic Region	Benick & Lohse 1974
		adjuvans Casey, 1910	Acrotona (s. str.) Thomson, 1859	149	ON	Gusarov 2003g
		fusiformis Casey, 1893	Achromata Casey, 1893	301	NY	Gusarov 2003g
		lividula Casey, 1910	Acrotona (s. str.) Thomson, 1859	147	OR	Gusarov 2003g
		nuptalis Casey, 1910	Dimetrota (s.str.) Mulsant & Rey, 1873	100	RI	Gusarov 2003g
Nehemitropia	lividipennis Mannerheim, 1830		Oxypoda Mannerheim, 1830	70	Palaearctic Region
Paraleptonia	pacei Klimaszewski in Klimaszewski & Winchester, 2002		Paraleptonia Klimaszewski in Klimaszewski & Winchester, 2002	47	BC	Klimaszewski & Winchester 2002
Parameotica	americana Bernhauer, 1907a		Atheta (Parameotica Ganglbauer, 1895) Thomson, 1858	386	n. nov. for Atheta sexualis Bernhauer, 1906 (nec Fauvel, 1899)
		californiana Pace, 1987	Atheta Thomson, 1858	436	n. nov. for Atheta sexualis Bernhauer, 1906 (nec Fauvel, 1899)
		sexualis Bernhauer, 1906	Atheta (Parameotica Ganglbauer, 1895) Thomson, 1858	339	CA
Philhygra	adjuncta Casey, 1910		Metaxya Mulsant & Rey, 1873	80	CA
		famula Casey, 1910	Metaxya Mulsant & Rey, 1873	81	CA
Philhygra	amniculivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		AK
Philhygra	angusticauda Bernhauer, 1909		Atheta (Metaxya Mulsant & Rey, 1873) Thomson, 1858	516	NH
Philhygra	aquivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		BC
Philhygra	aquulivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		CA
Philhygra	arenivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		OR
Philhygra	badeola Casey, 1911		Metaxya Mulsant & Rey, 1873	133	CA
Philhygra	benigna Casey, 1910		Hydrosmecta Thomson, 1858	88	CA
		jugalis Casey, 1910	Hydrosmecta Thomson, 1858	89	CA
Philhygra	botanicarum Muona, 1983		Philhygra Mulsant & Rey, 1873	57	Palaearctic Region	Muona 1983
Philhygra	bracata Casey, 1910		Metaxya Mulsant & Rey, 1873	79	NV
		discreta Casey, 1910	Metaxya Mulsant & Rey, 1873	79	NV
		triplicata Moore & Legner, 1975	Atheta (Brundinia Tottenham, 1949) Thomson, 1858	376	n. nov. for Atheta discreta Casey, 1910: 79 (nec Casey, 1893, Casey, 1910: 42)
Philhygra	clemens Casey, 1910		Noverota Casey, 1910	92	WI
		albanica Casey, 1911	Metaxya Mulsant & Rey, 1873	131	NY
		cephalica Bernhauer i.l.			PA
		elusa Casey, 1910	Metaxya Mulsant & Rey, 1873	78	IA
		impotens Casey, 1910	Metaxya Mulsant & Rey, 1873	81	RI
Philhygra	clivivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		CA
Philhygra	collinivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		BC
Philhygra	criddlei Casey, 1911		Metaxya Mulsant & Rey, 1873	137	MB
Philhygra	delectans Casey, 1910		Metaxya Mulsant & Rey, 1873	82	OR
Philhygra	discrepans Casey, 1910		Metaxya Mulsant & Rey, 1873	77	IA
		discordia Moore & Legner, 1975	Atheta (Brundinia Tottenham, 1949) Thomson, 1858	359	n. nov. for Atheta discrepans (Casey, 1910) (nec Sharp 1883)
Philhygra	falcifera Lohse in Lohse et al., 1990		Philhygra Mulsant & Rey, 1873	169	MB	Lohse et al. 1990
Philhygra	fatua Casey, 1910		Metaxya Mulsant & Rey, 1873	80	CA
Philhygra	finitima Casey, 1910		Noverota Casey, 1910	93	RI
		libens Casey, 1910	Atheta (Hilara Mulsant & Rey, 1873) Thomson, 1858	63	RI
		sejuncta Casey, 1910	Atheta (Hilara Mulsant & Rey, 1873) Thomson, 1858	64	RI
		validiceps Casey, 1910	Atheta (Hilara Mulsant & Rey, 1873) Thomson, 1858	62	RI
Philhygra	fluentivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		AB
Philhygra	fuscula Casey, 1906		Homalotusa Casey, 1906	341	ID
		acutiventris Bernhauer i.l.			ID
		pallida Casey, 1906	Homalotusa Casey, 1906	342	ID
Philhygra	helenica Casey, 1906		Homalotusa Casey, 1906	340	MT
		coloradensis Casey, 1911	Atheta (Homalotusa Casey, 1906) Thomson, 1858	81	CO
Philhygra	humivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		AK
Philhygra	iterans Casey, 1910		Atheta (s.str.) Thomson, 1858	20	NY
		perspicua Casey, 1911	Atheta (Delphota Casey, 1910) Thomson, 1858	85	NJ
Philhygra	junii Lohse in Lohse et al., 1990		Philhygra Mulsant & Rey, 1873	175	YT	Lohse et al. 1990
Philhygra	lacusculivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		AB
Philhygra	laevicollis Maeklin in Mannerheim, 1852		Homalota Mannerheim, 1830	306	AK	Lohse & Smetana 1985
		mordax Casey, 1911	Atheta (s.str.) Thomson, 1858	115	BC
		prognata Casey, 1911	Metaxya Mulsant & Rey, 1873	133	BC
Philhygra	leechi Lohse in Lohse et al., 1990		Philhygra Mulsant & Rey, 1873	173	NT	Lohse et al. 1990
Philhygra	limnivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		OR
Philhygra	lustrivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		NB
Philhygra	malleoides Lohse in Lohse et al., 1990		Philhygra Mulsant & Rey, 1873	171	NT	Lohse et al. 1990
Philhygra	manitobae Casey, 1911		Atheta (Homalotusa Casey, 1906) Thomson, 1858	81	MB
Philhygra	mateana Casey, 1911		Metaxya Mulsant & Rey, 1873	134	CA
Philhygra	mormon Casey, 1910		Atheta (Homalotusa Casey, 1906) Thomson, 1858	10	UT
Philhygra	muscivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		BC
Philhygra	nivivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		WA
Philhygra	nubivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		CA
Philhygra	nupera Casey, 1910		Atheta (s.str.) Thomson, 1858	40	NY
Philhygra	oregonina Casey, 1910		Atheta (Homalotusa Casey, 1906) Thomson, 1858	11	OR
Philhygra	oscitans Casey, 1911		Atheta (Delphota Casey, 1910) Thomson, 1858	88	CA
Philhygra	paludivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		AK
Philhygra	parallela Casey, 1906		Valenusa Casey, 1906	342	CA
Philhygra	palustris Kiesenwetter, 1844		Homalota Mannerheim, 1830	318	Palaearctic Region	Brundin 1944
		ithacana Casey, 1910	Atheta (Traumoecia Mulsant & Rey, 1873) Thomson, 1858	46	NY
		orbiceps Casey, 1910	Atheta (Traumoecia Mulsant & Rey, 1873) Thomson, 1858	47	IA
		subdola Casey, 1910	Atheta (Traumoecia Mulsant & Rey, 1873) Thomson, 1858	47	WI
		tenuicula Casey, 1911	Atheta (Traumoecia Mulsant & Rey, 1873) Thomson, 1858	118	MB
Philhygra	planivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		IL
Philhygra	proterminalis Bernhauer, 1907a		Atheta (Metaxya Mulsant & Rey, 1873) Thomson, 1858	383	PA
		bellula Casey, 1910	Metaxya Mulsant & Rey, 1873	77	RI
Philhygra	pseudoboreostiba Lohse in Lohse et al., 1990		Philhygra Mulsant & Rey, 1873	169	YT	Lohse et al. 1990
Philhygra	rarula Casey, 1910		Hydrosmecta Thomson, 1858	90	NY
	repens Casey, 1910		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	60	CA
Philhygra	ripicoloides Lohse in Lohse et al., 1990		Philhygra Mulsant & Rey, 1873	174	NT	Lohse et al. 1990
Philhygra	ripivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		NB
Philhygra	rostrifera Lohse in Lohse et al., 1990		Philhygra Mulsant & Rey, 1873	169	LB	Lohse et al. 1990
Philhygra	satanas Bernhauer, 1907a		Atheta (Metaxya Mulsant & Rey, 1873) Thomson, 1858	383	AB
Philhygra	saturata Casey, 1910		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	59	CA
Philhygra	scenica Casey, 1910		Noverota Casey, 1910	93	NY
Philhygra	silvivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		NT
Philhygra	stagnivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		AK
Philhygra	subfusca Casey, 1910		Metaxya Mulsant & Rey, 1873	83	CA
Philhygra	subpolaris Fenyes, 1909		Atheta (Metaxya Mulsant & Rey, 1873) Thomson, 1858	423	AZ
		deceptor Casey, 1910	Metaxya Mulsant & Rey, 1873	79	NM
Philhygra	tahoensis Casey, 1906		Homalotusa Casey, 1906	341	CA
Philhygra	terrivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		OR
Philhygra	vallivaga Gusarov, in prep.		Philhygra Mulsant & Rey, 1873		CA
Philhygra	varula Casey, 1911		Metaxya Mulsant & Rey, 1873	136	MB	Lohse et al. 1990 (as Philhygra stylifera)
		stylifera Lohse in Lohse et al., 1990	Philhygra Mulsant & Rey, 1873	174	PQ
Philhygra	wisconsinica Casey, 1910		Atheta (Homalotusa Casey, 1906) Thomson, 1858	12	WI
		awemeana Casey, 1911	Metaxya Mulsant & Rey, 1873	132	MB
		lacustrina Casey, 1910	Atheta (Homalotusa Casey, 1906) Thomson, 1858	11	WI
Pontomalota	opaca LeConte, 1863		Phytosus Curtis, 1838	28	CA	Ahn & Ashe 1992
		bakeri Bernhauer in Baker, 1912	Phytosus Curtis, 1838	170	CA	Ahn & Ashe 1992
		californica Casey, 1885	Pontomalota Casey, 1885	298	CA	Ahn & Ashe 1992
		luctuosa Casey, 1911	Pontomalota Casey, 1885	164	BC	Ahn & Ashe 1992
		nigriceps Casey, 1885	Pontomalota Casey, 1885	299	CA	Ahn & Ashe 1992
Pontomalota	terminalia Ahn & Ashe, 1992		Pontomalota Casey, 1885	356	CA	Ahn & Ashe 1992
Psammostiba	comparabilis Mäklin in Mannerheim, 1853		Homalota Mannerheim, 1830	181	AK	Gusarov 2003e
Psammostiba	kenaii Gusarov, 2003		Psammostiba Yosii & Sawada, 1976	28	AK	Gusarov 2003e
Sableta	infulata Casey, 1910		Sableta Casey, 1910	107	MS	Ashe & Gusarov in press
Schistacme	obtusa Notman, 1920		Schistacme Notman, 1920	712	FL
Schistoglossa	approximata Bernhauer, 1909		Atheta (Metaxya Mulsant & Rey, 1873) Thomson, 1858	516	MA
		flavicornis Bernhauer, 1909	Atheta (Amischa Thomson, 1858) Thomson, 1858	527	PA
Seeversiella	globicollis Bernhauer, 1907a		Atheta (Microdota Mulsant & Rey, 1873) Thomson, 1858	388	ON	Gusarov 2003a
		bispinosa Ashe, 1986	Seeversiella Ashe, 1986	503	AZ	Gusarov 2003a
Seeversiella	texana Gusarov, 2003a		Seeversiella Ashe, 1986	22	TX	Gusarov 2003a
Stethusa	dichroa Gravenhorst, 1802		Aleochara Gravenhorst, 1802	186	North America	Gusarov 2003f
		affluens Casey, 1910	Atheta (Stethusa Casey, 1910) Thomson, 1858	5	RI	Gusarov 2003f
		canonica Casey, 1910	Atheta (Stethusa Casey, 1910) Thomson, 1858	6	RI	Gusarov 2003f
		cernens Casey, 1911	Atheta (Stethusa Casey, 1910) Thomson, 1858	79	MS	Gusarov 2003f
		clarescans Casey, 1911	Atheta (Stethusa Casey, 1910) Thomson, 1858	77	NY	Gusarov 2003f
		cynica Casey, 1911	Atheta (Stethusa Casey, 1910) Thomson, 1858	78	RI	Gusarov 2003f
		galvestonica Casey, 1910	Atheta (Stethusa Casey, 1910) Thomson, 1858	6	TX	Gusarov 2003f
		irvingi Casey, 1910	Atheta (Stethusa Casey, 1910) Thomson, 1858	5	NY	Gusarov 2003f
		novella Casey, 1910	Dimetrota (s.str.) Mulsant & Rey, 1873	105	NY	Gusarov 2003f
		officiosa Casey, 1911	Atheta (Stethusa Casey, 1910) Thomson, 1858	79	NY	Gusarov 2003f
		sagax Casey, 1910	Atheta (Stethusa Casey, 1910) Thomson, 1858	7	VA	Gusarov 2003f
		sentiens Casey, 1910	Dimetrota (s.str.) Mulsant & Rey, 1873	105	KS	Gusarov 2003f
		sororella Casey, 1910	Atheta (Stethusa Casey, 1910) Thomson, 1858	7	NY	Gusarov 2003f
		tuta Casey, 1911	Atheta (Stethusa Casey, 1910) Thomson, 1858	80	NY	Gusarov 2003f
		videns Casey, 1911	Atheta (Stethusa Casey, 1910) Thomson, 1858	78	RI	Gusarov 2003f
Stethusa	klimschi Bernhauer, 1909		Atheta (s.str.) Thomson, 1858	523	LA	Gusarov 2003f
		sulcata Blatchley, 1910	Atheta Thomson, 1858	355	IN	Gusarov 2002d; Gusarov 2003f
Stethusa	spuriella Casey, 1910		Atheta (Stethusa Casey, 1910) Thomson, 1858	8	NY	Gusarov 2003f
		duplicata Fenyes, 1920	Atheta (s.str.) Thomson, 1858	213	n. nov. for Atheta unigranosa Bernhauer, 1909) (nec Bernhauer, 1908)	Gusarov 2003f
		macrops Notman, 1920	Atheta Thomson, 1858	725	FL	Gusarov 2003f
		subdebilis Casey, 1910	Atheta (Stethusa Casey, 1910) Thomson, 1858	7	MO	Gusarov 2003f
		unigranosa Bernhauer, 1909	Atheta (s.str.) Thomson, 1858	521	DE	Gusarov 2003f
Strigota	ambigua Erichson, 1839		Homalota Mannerheim, 1830	134	North America	Gusarov 2003g
		assueta Casey, 1910	Strigota Casey, 1910	178	MO	Gusarov 2003g
		brevipennis Notman, 1922	Anaduosternum Notman, 1922	107	NJ	Gusarov 2003g
		gnava Casey, 1910	Strigota Casey, 1910	177	NC	Gusarov 2003g
		inculta Casey, 1910	Strigota Casey, 1910	180	NV	Gusarov 2003g
		mediocris Casey, 1910	Strigota Casey, 1910	179	TX	Gusarov 2003g
		notmani Moore & Legner, 1975	Atheta (Anaduosternum Notman, 1922) Thomson, 1858	367	n. nov. for Atheta brevipennis (Notman, 1922) (nec Sahlberg, 1876)	Gusarov 2003g
		oppidana Casey, 1910	Strigota Casey, 1910	177	NY	Gusarov 2003g
		placata Casey, 1910	Strigota Casey, 1910	182	CA	Gusarov 2003g
		recta Casey, 1911	Strigota (s.str.) Casey, 1910	165	CT	Gusarov 2003g
		vapida Casey, 1910	Strigota Casey, 1910	179	NM	Gusarov 2003g
		verecunda Casey, 1910	Strigota Casey, 1910	178	IA	Gusarov 2003g
Strigota	perplexa Casey, 1910		Strigota Casey, 1910	180	CO
Strigota	seducens Casey, 1910		Strigota Casey, 1910	182	CA
		impiger Casey, 1910	Strigota Casey, 1910	181	WA
		intrudens Casey, 1910	Strigota Casey, 1910	181	CA
Strophogastra	penicillata Fenyes, 1921		Strophogastra Fenyes, 1921	21	MB
Tarphiota	fucicola Mäklin in Mannerheim, 1852		Tachyusa Erichson, 1837	75	BC	Ahn 1996
		debilicollis Casey, 1910	Tarphiota Casey, 1893	75	BC	Ahn 1996
		pallidipes Casey, 1893	Tarphiota Casey, 1893	333	CA	Ahn 1996
Tarphiota	geniculata Mäklin in Mannerheim, 1852		Homalota Mannerheim, 1830	308	AK	Ahn 1996
		litorina hirsutula Casey, 1910	Tarphiota Casey, 1893	75	CA	Ahn 1996
		insolita Casey, 1910	Tarphiota Casey, 1893	76	BC	Ahn 1996
		iota Casey, 1910	Tarphiota Casey, 1893	76	CA	Ahn 1996
		litorina Casey, 1910	Tarphiota Casey, 1893	75	CA	Ahn 1996
		seditiosa Casey, 1910	Tarphiota Casey, 1893	76	BC	Ahn 1996
Thamiaraea	americana Bernhauer, 1907a		Thamiaraea Thomson, 1858	401	LA
Thamiaraea	brittoni Casey, 1911		Sableta (Fusalia Casey, 1911) Casey, 1910	145	CT	Hoebeke 1988 (as Thamiaraea lira)
		lira Hoebeke, 1988	Thamiaraea Thomson, 1858	21	PA	Gusarov 2003g
		paralira Hoebeke, 1994	Thamiaraea Thomson, 1858	1	NY	Gusarov 2003g
Thinusa	maritima Casey, 1885		Phytosus Curtis, 1838	312	CA	Ahn 1997
		nigra Casey, 1911	Thinusa Casey, 1893	214	BC
		obscura Casey, 1906	Thinusa Casey, 1893	354	CA	Ahn 1997
Thinusa	fletcheri Casey, 1906		Thinusa Casey, 1893	353	BC	Ahn 1997
		divergens Casey, 1911	Thinusa Casey, 1893	213	BC	Ahn 1997
		fletcheri robustula Casey, 1911	Thinusa Casey, 1893	215	BC	Ahn 1997
Tomoglossa	arizonica Gusarov, 2002b		Tomoglossa Kraatz, 1856	14	FL	Gusarov 2002a
Tomoglossa	decora Casey, 1910		Noverota Casey, 1910	91	MS	Gusarov 2002a
Tomoglossa	floridana Gusarov, 2002b		Tomoglossa Kraatz, 1856	15	AZ	Gusarov 2002a
Tomoglossa	improvisa Casey, 1910		Noverota Casey, 1910	91	TX	Gusarov 2002a
Tomoglossa	ornatella Casey, 1910		Noverota Casey, 1910	90	NJ	Gusarov 2002a
Trichiusa	atra Casey, 1906		Trichiusa Casey, 1893	330	KS
Trichiusa	columbica Casey, 1911		Trichiusa Casey, 1893	76	BC
Trichiusa	compacta Casey, 1893		Trichiusa Casey, 1893	341	DC
		transversa Casey, 1906	Trichiusa Casey, 1893	327	OH
Trichiusa	convergens Casey, 1906		Trichiusa Casey, 1893	329	TX
Trichiusa	discreta Casey, 1906		Trichiusa Casey, 1893	331	ID
Trichiusa	hirsuta Casey, 1906		Trichiusa Casey, 1893	329	VA
Trichiusa	monticola Casey, 1906		Trichiusa Casey, 1893	328	ID
Trichiusa	parviceps Casey, 1906		Trichiusa Casey, 1893	328	OH
Trichiusa	parvicollis Casey, 1893		Trichiusa Casey, 1893	344	DE
Trichiusa	pilosa Casey, 1893		Trichiusa Casey, 1893	343	RI
Trichiusa	polita Casey, 1906		Trichiusa Casey, 1893	327	DC
Trichiusa	postica Casey, 1906		Trichiusa Casey, 1893	330	ON
Trichiusa	rigida Casey, 1906		Trichiusa Casey, 1893	331	TX
Trichiusa	robustula Casey, 1893		Trichiusa Casey, 1893	343	IA
Trichiusa	setigera Casey, 1893		Trichiusa Casey, 1893	342	NJ
Trichiusa	transversa Casey, 1906		Trichiusa Casey, 1893	327	OH
Trichiusa	ursina Notman, 1920		Trichiusa Casey, 1893	725	FL
Trichiusa	varicolor Casey, 1906		Trichiusa Casey, 1893	332	VA
Trichiusa	virginica Casey, 1906		Trichiusa Casey, 1893	330	VA
Tropimenelytron	americanum Gusarov, 2002e		Tropimenelytron Pace, 1983	12	IN	Gusarov 2002e
Tropimenelytron	californicum Gusarov, 2002e		Tropimenelytron Pace, 1983	17	CA	Gusarov 2002e
Tropimenelytron	robustum Gusarov, 2002e		Tropimenelytron Pace, 1983	14	OR	Gusarov 2002e

Species removed from Athetini

Species removed from Athetini		Page numbers in the original publications	The type locality states or provinces	Valid species names	Recent publications with a diagnoses	Recent publications with a confirmation of this synonymy	Correct tribe assignement
Species names	The genera (with subgenera) in which the species were originally described	Page numbers in the original publications	The type locality states or provinces	Valid species names	Recent publications with a diagnoses	Recent publications with a confirmation of this synonymy	Correct tribe assignement
canadensis Casey, 1910	Sableta (Canastota Casey, 1910) Casey, 1910	108	ON	Silusida marginella (Casey, 1893)	Ashe 1992	Ashe & Gusarov 2003	Homalotini Heer, 1839
cornicula Casey, 1911	Pseudota Casey, 1910	151	BC	Placusa vaga Casey, 1911	Klimaszewski et al. 2001	Gusarov 2003g	Placusini Mulsant & Rey 1872
densissima Benhauer, 1909	Atheta (Paradilacra Bernhauer, 1909) Thomson, 1858	517	CA	Paradilacra densissima Benhauer, 1909	Gusarov 2003g		Oxypodini Thomson, 1859
deserticola Casey, 1911	Paradilacra Bernhauer, 1909	131	NV	Paradilacra densissima Benhauer, 1909		Gusarov 2003g	Oxypodini Thomson, 1859
erebea Casey, 1911	Paradilacra Bernhauer, 1909	128	CA	Paradilacra densissima Benhauer, 1909		Gusarov 2003g	Oxypodini Thomson, 1859
filaria Casey, 1911	Sipalia (Sipaliella Casey, 1911) Mulsant & Rey, 1853	159	RI	Meotica pallens Redtenbacher, 1849	Muona 1991	Gusarov 2002c	Oxypodini Thomson, 1859
fortuita Casey, 1910	Colpodota Mulsant & Rey, 1873	164	MA	Oxypoda fortuita (Casey, 1910)			Oxypodini Thomson, 1859
glenorica Casey, 1910	Paradilacra Bernhauer, 1909	74	BC	Paradilacra densissima Benhauer, 1909		Gusarov 2003g	Oxypodini Thomson, 1859
ingratula Casey, 1910	Atheta (Phasmota Casey, 1910) Thomson, 1858	54	MS	Phasmota ingratula (Casey, 1910)			Oxypodini Thomson, 1859
luscitiosa Casey, 1910	Atheta (s. str.) Thomson, 1859	39	CA	Ocalea Erichson, 1837			Oxypodini Thomson, 1859
memnonia Casey, 1911	Paradilacra Bernhauer, 1909	130	CA	Paradilacra densissima Benhauer, 1909		Gusarov 2003g	Oxypodini Thomson, 1859
pallens Casey, 1893	Asthenesita Casey, 1893	366	FL	Asthenesita Casey, 1893			Homalotini Heer, 1839
pernix Casey, 1910	Dolosota (Microlia Casey, 1910) Casey, 1910	144	VA	Microlia silacea (Erichson, 1839)		Gusarov 2002b	Hoplandriini Casey, 1910
persola Casey, 1910	Paradilacra Bernhauer, 1909	72	CA	Paradilacra densissima Benhauer, 1909		Gusarov 2003g	Oxypodini Thomson, 1859
silacea Erichson, 1839	Homalota Mannerheim, 1830	120	North America	Microlia silacea (Erichson, 1839)	Gusarov 2002b		Hoplandriini Casey, 1910
sinistra Casey, 1911	Paradilacra Bernhauer, 1909	129	CA	Paradilacra densissima Benhauer, 1909		Gusarov 2003g	Oxypodini Thomson, 1859
sollemnis Casey, 1910	Ancillota Casey, 1910	165	MO	Oxypoda amica Casey, 1906		Gusarov 2003g	Oxypodini Thomson, 1859
subaequa Casey, 1911	Paradilacra Bernhauer, 1909	128	CA	Paradilacra densissima Benhauer, 1909		Gusarov 2003g	Oxypodini Thomson, 1859
symbolica Casey, 1911	Paradilacra Bernhauer, 1909	127	CA	Paradilacra densissima Benhauer, 1909		Gusarov 2003g	Oxypodini Thomson, 1859
uintana Casey, 1910	Paradilacra Bernhauer, 1909	73	UT	Paradilacra densissima Benhauer, 1909		Gusarov 2003g	Oxypodini Thomson, 1859
vulgata Casey, 1911	Paradilacra Bernhauer, 1909	130	CA	Paradilacra densissima Benhauer, 1909		Gusarov 2003g	Oxypodini Thomson, 1859
willametta Casey, 1910	Paradilacra Bernhauer, 1909	73	OR	Paradilacra densissima Benhauer, 1909		Gusarov 2003g	Oxypodini Thomson, 1859

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